Leaves with black dots
The following is a list of native or naturalised species which, at least some of the time, possess this spot-character.
The black dots are bacterial nodules in the following genera in the Rubiaceae: Pavetta, Psychotria and Sericanthe. Number of species: 4
Copyright: Mike Bingham, Annette Willemen, Bart Wursten, Petra Ballings and Mark Hyde, 2011-14
Bingham, M.G., Willemen, A., Wursten, B.T., Ballings, P. and Hyde, M.A.
. Flora of Zambia: List of species with each spot-character: Leaves with black dots.
http://www.zambiaflora.com/speciesdata/species-spot-display.php?species_id=749, retrieved 24 November 2014